Figure 5

Ordination diagram of species and environmental variables along the first two CCA axes. Herb layer – black triangles; shrub layer – black squares, tree layer – black circles; precip – annual sum of precipitation; elevat – elevation; temp – mean annual temperature; *statistically significant variables. For abbreviations of soil properties, see Table 1. Abbreviations (96 species, without 83 sporadic species): Aeg.pod - Aegopodium podagraria, All.pet - Alliaria petiolata, Aln.gluA - Alnus glutinosa (tree layer), Alo.pra - Alopecurus pratensis, Ane.nem - Anemone nemorosa, Ant.syl - Anthriscus sylvestris, Arr.ela - Arrhenatherum elatius, Art.vul - Artemisia vulgaris, Bid.fro - Bidens frondosa, Cal.epi - Calamagrostis epigeios, Cal.sep - Calystegia sepium, Car.acu - Carex acutiformis, Car.bri - Carex brizoides, Car.bue - Carex buekii, Car.hir - Carex hirta, Car.ela - Carex elata, Car.pra - Carex praecox, Che.maj - Chelidonium majus, Cir.arv - Cirsium arvense, Con.arv - Convolvulus arvensis, Cor.sanC - Cornus sanguinea (herb layer), Cra.monB - Crataegus monogyna (shrub layer), Cra.monC - Crataegus monogyna (herb layer), Dac.glo - Dactylis glomerata, Des.cae - Deschampsia caespitosa, Equ.arv - Equisetum arvense, Equ.pal - Equisteum palustre, Euo.eur - Euonymus europaeus, Eup.can - Eupatorium cannabinum, Fil.ulm - Filipendula ulmaria, Fil.vul - Filipendula vulgaris, Fra.ves - Fragaria vesca, Gal.bif - Galeopsis bifida, Gal.spe - Galeopsis speciosa, Gal.tet - Galeopsis tetrahit, Gal.apa - Galium aparine, Gal.bor - Galium boreale, Gal.mol - Galium mollugo, Gal.riv - Galium rivale, Gal.ver - Galium verum, Geu.urb - Geum urbanum, Her.sph - Heracleum sphondylium, Hol.lan - Holcus lanatus, Hum.lup - Humulus lupulus, Hyp.per - Hypericum perforatum, Imp.nol - Impatiens noli-tangere, Imp.par - Impatiens parviflora, Jun.effJuncus - effusus, Lam.mac - Lamium maculatum, Lat.pra - Lathyrus pratensis, Lyc.flo - Lychnis flos-cuculi, Lys.vul - Lysimachia vulgaris, Lyt.sal - Lythrum salicaria, Men.lon - Mentha longifolia, Myo.pal - Myosotis palustris, Myo.aqu - Myososton aquaticum, Pad.aviA - Padus avium (tree layer), Pet.hyb - Petasites hybridus, Pha.aru - Phalaris arundinacea, Phr.aus - Phragmites australis, Poa.pra - Poa pratensis, Poa.tri - Poa trivialis, Pol.amp - Polygonum amphibium f. terrestre, Pol.hyd - Polygonum hydropiper, Pot.rep - Potentilla reptans, Pru.spiC - Prunus spinosa (herb layer), Que.robA - Quercus robur (tree layer), Ran.acr - Ranunculus acris, Rub.caeC - Rubus caesius (herb layer), Rub.ida -Rubus idaeus, Rub.fruC - Rubus fruticosus agg. (herb layer), Rum.ace - Rumex acetosa, Rum.thy - Rumex thyrsiflorus, Sal.albA - Salix alba (tree layer), Sal.fraA –Salix fragilis (tree layer), Sam.nigC - Sambucus nigra (herb layer), San.off - Sanguisorba officinalis, Sci.syl - Scirpus sylvaticus, Scr.nod - Scrophularia nodosa, Scu.gal - Scutellaria galericulata, Sel.cer - Selinum carvifolia, Sil.alb - Silene alba, Sol.dul - Solanum dulcamara, Sol.gig - Solidago gigantea, Sta.pal - Stachys palustris, Sta.syl - Stachys sylvatica, Ste.gra - Stellaria graminea, Ste.med - Stellaria media, Sym.off - Symphytum officinale, Tan.vul - Tanacetum vulgare, Tha.luc - Thalictrum lucidum, Urt.dio - Urtica dioica, Ver.lon - Veronica longifolia, Vib.opu - Viburnum opulusC (herb layer), Vic.cra - Vicia cracca.