Author Correction: Universal DNA methylation age across mammalian tissues

1. These authors contributed equally: A. T. Lu, Z. Fei, K. Raj, S. Horvath.

Authors and Affiliations

1. Department of Human Genetics, David Geffen School of Medicine, University of California, Los Angeles, Los Angeles, CA, USA

   A. T. Lu, A. Haghani, J. Zhang, A. R. Lim & S. Horvath

2. Altos Labs, San Diego Institute of Science, San Diego, CA, USA

   A. T. Lu, A. Haghani, Q. Yan, J. C. Izpisua Belmonte, P. Reddy & S. Horvath

3. Department of Biostatistics, Fielding School of Public Health, University of California, Los Angeles, Los Angeles, CA, USA

   Z. Fei, J. A. Zoller, C. Z. Li, M. Yao & S. Horvath

4. Department of Statistics, University of California, Riverside, Riverside, CA, USA

   Z. Fei

5. Zoological SeaWorld Parks and Entertainment, Orlando, FL, USA

   T. R. Robeck

6. Altos Labs, Cambridge Institute of Science, Cambridge, UK

   R. Lowe & K. Raj

7. Bioinformatics Interdepartmental Program, University of California, Los Angeles, CA, USA

   H. Vu, A. Arneson, S. B. Kwon & J. Ernst

8. Department of Biological Chemistry, University of California, Los Angeles, Los Angeles, CA, USA

   H. Vu, A. Arneson, S. B. Kwon & J. Ernst

9. Department of Biology, University of Rochester, Rochester, NY, USA

   J. Ablaeva

10. Department of Neuroscience, Peter O’Donnell Jr. Brain Institute, University of Texas Southwestern Medical Center, Dallas, TX, USA

    V. A. Acosta-Rodriguez, C. B. Green & J. S. Takahashi

11. Department of Biology, University of Maryland, College Park, MD, USA

    D. M. Adams & G. S. Wilkinson

12. Loro Parque Fundacion, Puerto de la Cruz, Spain

    J. Almunia

13. Department of Biology, University of Kentucky, Lexington, KY, USA

    A. Aloysius & A. W. Seifert

14. Division of Cardiology, Department of Internal Medicine, David Geffen School of Medicine, University of California, Los Angeles, Los Angeles, CA, USA

    R. Ardehali, N. B. Nguyen & P. Zhao

15. Marine Mammal Institute, Oregon State University, Newport, OR, USA

    C. S. Baker & E. K. Bors

16. School of Science and Technology, Clifton Campus, Nottingham Trent University, Nottingham, UK

    G. Banks

17. School of Life and Environmental Sciences, the University of Sydney, Sydney, New South Wales, Australia

    K. Belov & C. J. Hogg

18. Department of Zoology and Entomology, University of Pretoria, Hatfield, South Africa

    N. C. Bennett & D. W. Hart

19. Busch Gardens Tampa, Tampa, FL, USA

    P. Black, M. C. Spriggs & N. Stedman

20. Department of Ecology and Evolutionary Biology, University of California, Los Angeles, Los Angeles, CA, USA

    D. T. Blumstein

21. Rocky Mountain Biological Laboratory, Crested Butte, CO, USA

    D. T. Blumstein

22. Altius Institute for Biomedical Sciences, Seattle, WA, USA

    C. E. Breeze

23. Epigenetic Clock Development Foundation, Los Angeles, CA, USA

    R. T. Brooke

24. Center for Species Survival, Smithsonian Conservation Biology Institute, Front Royal, VA, USA

    J. L. Brown

25. Department of Evolution, Ecology and Organismal Biology, The Ohio State University, Columbus, OH, USA

    G. G. Carter

26. AgResearch, Invermay Agricultural Centre, Mosgiel, New Zealand

    A. Caulton & S. M. Clarke

27. Department of Biochemistry, University of Otago, Dunedin, New Zealand

    A. Caulton

28. Gulf World, Dolphin Company, Panama City Beach, FL, USA

    J. M. Cavin

29. School of Veterinary Medicine and Science, University of Nottingham, Nottingham, UK

    L. Chakrabarti

30. Department of Pathology, Microbiology and Immunology, School of Medicine, University of South Carolina, Columbia, SC, USA

    I. Chatzistamou

31. Department of Pharmacology, Addiction Science and Toxicology, the University of Tennessee Health Science Center, Memphis, TN, USA

    H. Chen

32. Medical Informatics, David Geffen School of Medicine, University of California, Los Angeles, Los Angeles, CA, USA

    K. Cheng

33. Biochemistry Research Institute of La Plata, Histology and Pathology, School of Medicine, University of La Plata, La Plata, Argentina

    P. Chiavellini, R. G. Goya & M. Lehmann

34. Center for Neurobehavioral Genetics, Semel Institute for Neuroscience and Human Behavior, Department of Psychiatry and Biobehavioral Sciences, David Geffen School of Medicine, University of California, Los Angeles, Los Angeles, CA, USA

    O. W. Choi, J. DeYoung & A. J. Jasinska

35. Department of Anatomy and Neurobiology, Northeast Ohio Medical University, Rootstown, OH, USA

    L. N. Cooper

36. Department of Environmental and Life Sciences, Trent University, Peterborough, Ontario, Canada

    M. L. Cossette

37. Taronga Institute of Science and Learning, Taronga Conservation Society Australia, Mosman, New South Wales, Australia

    J. Day & J. K. O’Brien

38. SeaWorld of Florida, Orlando, FL, USA

    S. DiRocco, C. Erlacher-Reid, D. M. Lindemann & L. Staggs

39. Zoological Operations, SeaWorld Parks and Entertainment, Orlando, FL, USA

    C. Dold & G. A. Montano

40. Duke Lemur Center, Durham, NC, USA

    E. E. Ehmke & M. Simmons

41. Conservation Biology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration, Seattle, WA, USA

    C. K. Emmons, M. B. Hanson & K. M. Parsons

42. Departments of Biology and Medicine, University of Rochester, Rochester, NY, USA

    S. Emmrich, V. Gorbunova, N. Macoretta, E. Rydkina, A. Seluanov, M. Takasugi, Z. Zhang & Y. Zhao

43. Altos Labs, San Francisco, CA, USA

    E. Erbay

44. SeaWorld Orlando, Orlando, FL, USA

    C. Erlacher-Reid

45. School of Biological and Behavioural Sciences, Queen Mary University of London, London, UK

    C. G. Faulkes

46. Fisheries and Oceans Canada, Freshwater Institute, Winnipeg, Manitoba, Canada

    S. H. Ferguson

47. Department of Biological Sciences, University of Manitoba, Winnipeg, Manitoba, Canada

    S. H. Ferguson

48. Department of Population Health and Reproduction, University of California, Davis School of Veterinary Medicine, Davis, CA, USA

    C. J. Finno & E. N. Hales

49. Mystic Aquarium, Mystic, CT, USA

    J. E. Flower

50. Universite de Lyon, Universite Lyon 1, CNRS, Laboratoire de Biometrie et Biologie Evolutive, Villeurbanne, France

    J. M. Gaillard, J. F. Lemaitre & B. Rey

51. Greenland Institute of Natural Resources, Nuuk, Greenland

    E. Garde

52. Evolution and Ecology Research Centre, School of Biological, Earth and Environmental Sciences, UNSW Sydney, Sydney, New South Wales, Australia

    L. Gerber

53. Division of Genetics, Department of Medicine, Brigham and Women’s Hospital, Harvard Medical School, Boston, MA, USA

    V. N. Gladyshev, S. G. Lee, J. R. Poganik, A. V. Shindyapina & B. Zhang

54. Applied Translational Genetics Group, School of Biological Sciences, Centre for Brain Research, the University of Auckland, Auckland, New Zealand

    M. J. Grant, P. Narayan & R. G. Snell

55. Vancouver Aquarium, Vancouver, British Columbia, Canada

    M. Haulena

56. SeaWorld of California, San Diego, CA, USA

    K. Herrick, J. Russell & T. Schmitt

57. Cancer Genetics and Comparative Genomics Branch, National Human Genome Research Institute, National Institutes of Health, Bethesda, MD, USA

    A. N. Hogan, E. A. Ostrander & J. Plassais

58. Department of Anatomy, University of Otago, Dunedin, New Zealand

    T. A. Hore & V. J. Sugrue

59. Division of Human Genetics, Department of Pediatrics, University at Buffalo, Buffalo, NY, USA

    T. Huang & J. Slone

60. Division of Genetics and Metabolism, Oishei Children’s Hospital, Buffalo, NY, USA

    T. Huang

61. School of Biological Sciences, University of Bristol, Bristol, UK

    G. Jones

62. Norwegian Orca Survey, Andenes, Norway

    E. Jourdain

63. Mother Infant Research Institute, Tufts Medical Center, Boston, MA, USA

    O. Kashpur, P. O’Tierney Ginn & M. C. Wallingford

64. Yuvan Research, Mountain View, CA, USA

    H. Katcher & A. Sanghavi

65. Kamogawa Sea World, Kamogawa, Japan

    E. Katsumata

66. Peromyscus Genetic Stock Center, University of South Carolina, Columbia, SC, USA

    V. Kaza & H. Kiaris

67. Department of Drug Discovery and Biomedical Sciences, College of Pharmacy, University of South Carolina, Columbia, SC, USA

    H. Kiaris, A. Naderi & E. Soltanmaohammadi

68. Edwin S.H. Leong Healthy Aging Program, Centre for Molecular Medicine and Therapeutics, University of British Columbia, Vancouver, British Columbia, Canada

    M. S. Kobor

69. Institute of Animal Reproduction and Food Research of the Polish Academy of Sciences, Olsztyn, Poland

    P. Kordowitzki

70. Institute for Veterinary Medicine, Nicolaus Copernicus University, Torun, Poland

    P. Kordowitzki

71. LGL Limited, King City, Ontario, Canada

    W. R. Koski

72. Evolutionary Genetics Group, Department of Evolutionary Anthropology, University of Zurich, Zurich, Switzerland

    M. Krützen & K. J. Peters

73. Department of Ecology and Evolutionary Biology, UCLA, Los Angeles, CA, USA

    B. Larison, G. M. Pinho, K. E. Sears, I. Sinha & R. K. Wayne

74. Center for Tropical Research, Institute for the Environment and Sustainability, UCLA, Los Angeles, CA, USA

    B. Larison

75. Department of Neurology, David Geffen School of Medicine, University of California, Los Angeles, Los Angeles, CA, USA

    A. J. Levine, K. C. Paul & C. K. Williams

76. Texas Pregnancy and Life-course Health Center, Southwest National Primate Research Center, San Antonio, TX, USA

    C. Li & P. W. Nathanielsz

77. Department of Animal Science, College of Agriculture and Natural Resources, Laramie, WY, USA

    C. Li & P. W. Nathanielsz

78. Technology Center for Genomics and Bioinformatics, Department of Pathology and Laboratory Medicine, University of California, Los Angeles, Los Angeles, CA, USA

    X. Li

79. Centre for Molecular Medicine and Therapeutics, BC Children’s Hospital Research Institute, University of British Columbia, Vancouver, British Columbia, Canada

    D. T. S. Lin

80. Institute of Ecology and Evolution, School of Biological Sciences, University of Edinburgh, Edinburgh, UK

    T. J. Little

81. White Oak Conservation, Yulee, FL, USA

    D. Maddox

82. North Gulf Oceanic Society, Homer, AK, USA

    C. O. Matkin

83. Translational Gerontology Branch, National Institute on Aging Intramural Research Program, National Institutes of Health, Baltimore, MD, USA

    J. A. Mattison

84. ABS Global, DeForest, WI, USA

    M. McClure

85. Marineland of Canada, Niagara Falls, Ontario, Canada

    J. Mergl

86. Biomedical and Genomic Research Group, Department of Animal and Dairy Sciences, University of Wisconsin—Madison, Madison, WI, USA

    J. J. Meudt & D. Shanmuganayagam

87. Department of Preventive Medicine, University of Tennessee Health Science Center, College of Medicine, Memphis, TN, USA

    K. Mozhui

88. Department of Genetics, Genomics and Informatics, University of Tennessee Health Science Center, College of Medicine, Memphis, TN, USA

    K. Mozhui & R. W. Williams

89. Louis Calder Center—Biological Field Station, Department of Biological Sciences, Fordham University, Armonk, NY, USA

    J. Munshi-South

90. Museum fur Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Berlin, Germany

    M. Nagy

91. Institute of Molecular Biology, Mainz, Germany

    C. Niehrs & L. Schomacher

92. Division of Molecular Embryology, DKFZ-ZMBH Alliance, Heidelberg, Germany

    C. Niehrs

93. Department of Obstetrics and Gynecology, Tufts University School of Medicine, Boston, MA, USA

    P. O’Tierney Ginn

94. Cancer Research UK Cambridge Institute, University of Cambridge, Cambridge, UK

    D. T. Odom

95. Division of Regulatory Genomics and Cancer Evolution, Deutsches Krebsforschungszentrum, Heidelberg, Germany

    D. T. Odom

96. Department of Psychology, Cornell University, Ithaca, NY, USA

    A. G. Ophir & L. L. Sailer

97. SeaWorld of Texas, San Antonio, TX, USA

    S. Osborn

98. Department of Molecular Cell and Developmental Biology, University of California, Los Angeles, Los Angeles, CA, USA

    M. Pellegrini, K. E. Sears & M. J. Thompson

99. School of Earth, Atmospheric and Life Sciences, University of Wollongong, Wollongong, Australia

    K. J. Peters

100. Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, UK

     A. B. Pedersen

101. Department of Animal Science, University of Nebraska, Lincoln, NE, USA

     J. L. Petersen & M. L. Spangler

102. Mammal Research Institute, Department of Zoology and Entomology, University of Pretoria, Hatfield, South Africa

     D. W. Pietersen

103. Department of Biology, College of Arts and Science, Adelphi University, Garden City, NY, USA

     N. A. Prado

104. Salk Institute for Biological Studies, La Jolla, CA, USA

     P. Reddy

105. Department of Epidemiology, UCLA Fielding School of Public Health, Los Angeles, CA, USA

     B. R. Ritz

106. Department of Environmental Health Sciences, UCLA Fielding School of Public Health, Los Angeles, CA, USA

     B. R. Ritz

107. Department of Neurology, UCLA David Geffen School of Medicine, Los Angeles, CA, USA

     B. R. Ritz

108. Center for Coastal Studies, Provincetown, MA, USA

     J. Robbins

109. Miami Seaquarium, Miami, FL, USA

     M. Rodriguez

110. The Sam and Ann Barshop Institute for Longevity and Aging Studies and Department of Molecular Medicine, UT Health San Antonio and the Geriatric Research Education and Clinical Center, South Texas Veterans Healthcare System, San Antonio, TX, USA

     A. B. Salmon

111. Department of Radiology, University of Illinois at Chicago, Chicago, IL, USA

     K. M. Schachtschneider & L. B. Schook

112. Department of Biochemistry and Molecular Genetics, University of Illinois at Chicago, Chicago, IL, USA

     K. M. Schachtschneider

113. National Center for Supercomputing Applications, University of Illinois at Urbana-Champaign, Urbana, IL, USA

     K. M. Schachtschneider

114. College of Agriculture, Missouri State University, Springfield, MO, USA

     D. Schmitt

115. Department of Animal Sciences, University of Illinois at Urbana-Champaign, Champaign, IL, USA

     L. B. Schook

116. Department of Forensic Science, Environmental and Life Sciences, Trent University, Peterborough, Ontario, Canada

     A. B. A. Shafer

117. Department of Surgery, University of Wisconsin School of Medicine and Public Health, Madison, WI, USA

     D. Shanmuganayagam

118. Shobhaben Pratapbhai Patel School of Pharmacy and Technology Management, SVKM’S NMIMS University, Mumbai, India

     K. Singh

119. Species Preservation Laboratory, SeaWorld San Diego, San Diego, CA, USA

     K. J. Steinman

120. Biology Department, Acadia University, Wolfville, Nova Scotia, Canada

     D. T. Stewart

121. Department of Pathobiology and Population Sciences, Royal Veterinary College, Hatfield, UK

     B. Szladovits

122. Howard Hughes Medical Institute, Department of Neuroscience, University of Texas Southwestern Medical Center, Dallas, TX, USA

     J. S. Takahashi

123. School of Biology and Environmental Science, University College Dublin, Dublin, Ireland

     E. C. Teeling

124. John G. Shedd Aquarium, Chicago, IL, USA

     B. Van Bonn

125. School of Biology, the University of St Andrews, Fife, UK

     S. C. Vernes

126. Neurogenetics of Vocal Communication Group, Max Planck Institute for Psycholinguistics, Nijmegen, the Netherlands

     S. C. Vernes

127. Blizard Institute, Faculty of Medicine and Dentistry, Queen Mary University of London, London, UK

     D. Villar

128. Department of Pathology and Laboratory Medicine, David Geffen School of Medicine at UCLA, Los Angeles, CA, USA

     H. V. Vinters

129. Division of Obstetrics and Gynecology, Tufts University School of Medicine, Boston, MA, USA

     M. C. Wallingford

130. Center for Neurobehavioral Genetics, Jane and Terry Semel Institute for Neuroscience and Human Behavior, University of California, Los Angeles, Los Angeles, CA, USA

     N. Wang & X. W. Yang

131. Department of Psychiatry and Biobehavioral Sciences, David Geffen School of Medicine at UCLA, Los Angeles, CA, USA

     N. Wang & X. W. Yang

132. Fisheries and Oceans Canada, Winnipeg, Manitoba, Canada

     B. G. Young

133. Eli and Edythe Broad Center of Regenerative Medicine and Stem Cell Research, University of California, Los Angeles, CA, USA

     P. Zhao

134. Center for Computational and Genomic Medicine, Children’s Hospital of Philadelphia, Philadelphia, PA, USA

     W. Zhou

135. Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, PA, USA

     W. Zhou

136. Department of Mathematics and Technology, University of Applied Sciences Koblenz, Koblenz, Germany

     J. Zimmermann

Corresponding author

Correspondence to S. Horvath.

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